Do "=" chars in request parameters need to be URL-encoded?

If I have a request parameter value that represents an LDAP
distinguished name, like "low=abc,middle=def,high=ghi", am I required to
URL-encode the "=" characters, so I would have to use
"low%3Dabc,middle%3Ddef,high%3Dghi" instead?

I mostly wonder what the specification states (and where in the
specification it would say this, and which specification).  I noticed
that Tomcat 4.1.18 lets me get away with not encoding them, but a very
old iPlanet installation gets confused if I don't encode them.

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