[dinosaur] GSA abstracts + Gorynychus + Russian Mesozoic mammals + "Cretaceous dicynodont" a marsupial + more

Ben Creisler
[email protected]


Some recent items and papers not yet mentioned (mainly non-dino stuff):

Geological Society of America Annual Meeting Sept. 22-25, 2019 abstracts
are now posted online. Presumably, there is an embargo on discussing the
content until the actual presentations and posters are officially made
public.

Search terms for dinosaur, pterosaur, etc., will bring up abstracts.

https://urldefense.proofpoint.com/v2/url?u=https-3A__gsa.confex.com_gsa_2019AM_meetingapp.cgi_Search_0-3Fsort-3DRelevance-26size-3D10-26page-3D1&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=WE9zClPKmPIBErFmto7yQd_Y0zM-axXkq4hGzfIA3Ww&e= 


https://urldefense.proofpoint.com/v2/url?u=https-3A__gsa.confex.com_gsa_2019AM_webprogram_start.html&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=uR3ZynOhfcUaYLBlIRvYOIhL8ImaYHE-kl9wQ1EnrDw&e= 

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Previously posted in Russian version, now in English translation:

A. O. Averianov & A. V. Lopatin (2019)
Dinosaur Fossils from the Upper Cretaceous of Crimea.
Paleontological Journal 53(4): 398-410
 DOI: https://urldefense.proofpoint.com/v2/url?u=https-3A__doi.org_10.1134_S0031030119040026&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=t_LZgCGj2M0H3urYepx9thOmKcFUJnKRh_1rkDwLcho&e= 
https://urldefense.proofpoint.com/v2/url?u=https-3A__link.springer.com_article_10.1134_S0031030119040026&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=wIt8px0EyllrDOcwfDfLyevCyKRVoODBjL18oHaL7S4&e= 


A new study of the holotype of Riabininohadros weberae from the Upper
Cretaceous (upper Maastrichtian) of Crimea (Besh-Kosh) reveals previously
unknown fragments of femur, astragalus, and calcaneus. This taxon is
characterized by a complex of primitive characters found in Iguanodontia
and basal Ornithischia and its phylogenetic position can be defined as
Styracosterna indet. The second dinosaur record from Crimea (Aleshino) is
represented by a fragmentary skeleton, including cervical and dorsal
vertebrae. It may belong to a derived iguanodontian or a primitive
hadrosauroid. Thus, at least two species of dinosaurs coexisted in the
Maastrichtian of Crimea.

==

Previously posted in Russian version, here in English translation:

M. A. Rogov, N. G. Zverkov, V. A. Zakharov & M. S. Arkhangelsky (2019)
Marine Reptiles and Climates of the Jurassic and Cretaceous of Siberia.
Stratigraphy and Geological Correlation 27(4):  398-423
DOI: https://urldefense.proofpoint.com/v2/url?u=https-3A__doi.org_10.1134_S0869593819040051&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=TEg6Gq-lOXykBa8Qn1yoJGopT_XeKaMzC815Lyu5oXc&e= 
https://urldefense.proofpoint.com/v2/url?u=https-3A__link.springer.com_article_10.1134_S0869593819040051&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=M6JxM-4C9bbyDgDDWYMnjSWyI3VA-0MuMxinEKVWjlI&e= 

Free pdf:
https://urldefense.proofpoint.com/v2/url?u=https-3A__www.researchgate.net_publication_334882493-5FMarine-5FReptiles-5Fand-5FClimates-5Fof-5Fthe-5FJurassic-5Fand-5FCretaceous-5Fof-5FSiberia&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=ZvqAzsajLaOuSNtOn6PXfgwZX2kbSXiEL2Tdqj27qbE&e= 

All current data on the Jurassic and Cretaceous climates of Siberia based
on isotope, paleontological, and lithological proxies are summarized. The
late Pliensbachian cooling episode, early Toarcian warming, promptly
replaced by long-term Middle Jurassic cooling at the end of the Toarcian,
and a long-term warm interval in the Late Jurassic are clearly recorded.
From the end of the Ryazanian, a gradual cooling episode began, which
apparently continued throughout the Early Cretaceous except for a brief
warming episode in the early Aptian. At the beginning of the Late
Cretaceous, the climate became warmer; the peak of warming is recorded at
the Cenomanian–Turonian boundary. Then, the middle–late Turonian was marked
by a relatively cold episode. Later, in the Coniacian–Campanian, the
climate warmed again, but at the end of the Campanian another cooling
episode occurred. New findings of marine reptiles are described from the
Toarcian, Kimmeridgian, Volgian, and Santonian-Campanian deposits of north
of Eastern Siberia. All existing records of marine reptiles known from the
Jurassic and Cretaceous of Siberia are revised, and all the findings (from
51 localities) are positioned in relation to paleolatitudes. It is
established that the majority of occurrences of these fossils were within
the polar paleolatitudes (70°–87°). We found no direct correlation between
climate fluctuations and the distribution of these organisms. Taking into
account the newest data showing that representatives of the majority of
Jurassic and Cretaceous large groups of marines reptiles were able to
maintain a more or less constant body temperature and were also able to
undertake large-scale seasonal migrations, it is reasonable to be cautious
in interpreting the presence of remains of these animals as indicators of a
warm climate.

=====

This article was posted earlier in the Russian version, now in English
translation with free pdf link:

Gorynychus sundyrensis sp. nov.

Yu. A. Suchkova & V. K. Golubev (2019)
A New Permian Therocephalian (Therocephalia, Theromorpha) from the Sundyr
Assemblage of Eastern Europe.
Paleontological Journal 53(4): 411-417
DOI: https://urldefense.proofpoint.com/v2/url?u=https-3A__doi.org_10.1134_S0031030119040117&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=ayKj3xFpx-g42PxqQtOLqFN9f2jysfne4vXz0ZG29iQ&e= 
https://urldefense.proofpoint.com/v2/url?u=https-3A__link.springer.com_article_10.1134_S0031030119040117&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=sVhk_LNO79U6U36xVlAD6TZ-aKUzLSagT2DUrHBhWgA&e= 

Free pdf:
https://urldefense.proofpoint.com/v2/url?u=https-3A__www.researchgate.net_publication_334807205-5FA-5FNew-5FPermian-5FTherocephalian-5FTherocephalia-5FTheromorpha-5Ffrom-5Fthe-5FSundyr-5FAssemblage-5Fof-5FEastern-5FEurope&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=7GW6KOfDlBZT5wSzMZ4TmRQpResOVCCGJc4fjN5nE48&e= 

Lycosuchid therocephalian Gorynychus sundyrensis sp. nov. is described
based on material from the middle Permian locality of Sundyr-1, Mari El
Republic, Russia. Within the Sundyr tetrapod community, G. sundyrensis
occupied the niche of a large scavenger.

====


Free pdf:

Alexey V. Lopatin, Alexander Averianov & S. V. Ivantsov (2019)
Two new localities of Mesozoic mammals in Russia (Krasnoyarsk territory,
Lower Cretaceous).
Doklady Akademii Nauk 487(4): 414-417 (in Russian)
DOI: 10.31857/S0869-56524874414-417
https://urldefense.proofpoint.com/v2/url?u=https-3A__journals.eco-2Dvector.com_0869-2D5652_article_view_15833&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=YUwjitQVeBrpM3Kut7L5H-tGeORbqe_ps3OTYgrPvFw&e= 

Free pdf:
https://urldefense.proofpoint.com/v2/url?u=https-3A__journals.eco-2Dvector.com_0869-2D5652_article_view_15833_12456&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=whuOjZoZjkNRYbyP48uoqQoqwi-5FQmTkHfjwm0T9qk&e= 


Two new localities of Mesozoic mammals have been discovered: Bol’shoi Ilek
and Berezovaya River (Russia, Krasnoyarsk Territory, Lower Cretaceous, Ilek
Formation). Bol’shoi Ilek locality yields an edentulous fragment of the
maxillary of Docodonta indet. A fragment of dentary without teeth
attributed to Mammalia indet. (presumably eutriconodontan or
symmetrodontan) is presented in the Berezovaya River locality. New
localities fill the geographical gap between previously known mammalian
localities of Ilek Formation in the basins of Kiya and Bol’shoi Kemchug
rivers.

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Free pdf:

Julia Lachner, Florian Ehrlich, Veronika Mlitz, Marcela Hermann, Lorenzo
Alibardi, Erwin Tschachler & Leopold Eckhart (2019)
Immunolocalization and phylogenetic profiling of the feather protein with
the highest cysteine content.
Protoplasma 256(5): 1257--1265
DOI: https://urldefense.proofpoint.com/v2/url?u=https-3A__doi.org_10.1007_s00709-2D019-2D01381-2D3&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=tbhMsa9_VP-i4UTFTW5tEknh686u-u2LtAKxqixoXYI&e= 
https://urldefense.proofpoint.com/v2/url?u=https-3A__link.springer.com_article_10.1007_s00709-2D019-2D01381-2D3&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=DM3Hyad3bQ7nHyboDMHj_VN4jNc5kzDW5tyJQ8LeCmM&e= 

Free pdf:
https://urldefense.proofpoint.com/v2/url?u=https-3A__link.springer.com_content_pdf_10.1007-252Fs00709-2D019-2D01381-2D3.pdf&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=4f5T2HvWQs5plDhv13Ug72_wUsaEi338_3nM5XqH2vk&e= 



Feathers are the most complex skin appendages of vertebrates. Mature
feathers consist of interconnected dead keratinocytes that are filled with
heavily cross-linked proteins. Although the molecular architecture
determines essential functions of feathers, only few feather proteins have
been characterized with regard to their amino acid sequences and evolution.
Here, we identify Epidermal Differentiation protein containing DPCC Motifs
(EDDM) as a cysteine-rich protein that has co-evolved with other feather
proteins. The EDDM gene is located within the avian epidermal
differentiation complex (EDC), a cluster of genes that has originated and
diversified in amniotes. EDDM shares the exon-intron organization with EDC
genes of other amniotes, including humans, and a gene encoding an EDDM-like
protein is present in crocodilians, suggesting that avian EDDM arose by
sequence modification of an epidermal differentiation gene present in a
common ancestor of archosaurs. The EDDM protein contains multiple sequence
repeats and a higher number of cysteine residues than any other protein
encoded in the EDC. Immunohistochemical analysis of chicken skin and skin
appendages showed expression of EDDM in barb and barbules of feathers as
well as in the subperiderm on embryonic scutate scales. These results
suggest that the diversification and differential expression of EDDM,
besides other EDC genes, was instrumental in facilitating the evolution of
the most complex molecular architecture of feathers.

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Free pdf:

Lílian P. Bergqvist, Paulo Victor Luiz G.C. Pereira, Alessandra S. Machado,
Mariela C. de Castro, Luiza B. Melki & Ricardo T. Lopes (2019)
Osteoderm microstructure of Riostegotherium yanei, the oldest Xenarthra.
Anais da Academia Brasileira de Ciências 91, suppl. 2: e20181290
doi: https://urldefense.proofpoint.com/v2/url?u=http-3A__dx.doi.org_10.1590_0001-2D3765201920181290&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=omXsp5GWW0ieHq0Y5pAKksW8QCwau0MpiRBFrQmu94c&e= 
https://urldefense.proofpoint.com/v2/url?u=http-3A__www.scielo.br_scielo.php-3Fscript-3Dsci-5Fabstract-26pid-3DS0001-2D37652019000400518-26lng-3Den-26nrm-3Diso-26tlng-3Den&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=Vx5pJfqD05IKmfE5TGg-pX2IoXoZdSpkZgjC-88yyAQ&e= 

Free pdf:
https://urldefense.proofpoint.com/v2/url?u=http-3A__www.scielo.br_pdf_aabc_v91s2_0001-2D3765-2Daabc-2D91-2Ds2-2De20181290.pdf&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=vKBrYnJAJhWX85Xt7k3P5TZ8NxqqQx0fiP3KpX9sU_g&e= 


Riostegotherium yanei from the Itaboraí Basin, Brazil, is the oldest known
Xenarthra. This paper aims to describe the internal morphology of the
osteoderms of Riostegotherium yanei from the perspective of histology and
micro-CT approaches, expanding the available data on cingulate osteoderm
microstructure. Seven osteoderms of R. yanei were used for the internal
microstructure description and eight of Dasypus novemcinctus for
comparison. The osteoderms of Riostegotherium yanei lacks the diploë-like
structure typical of glyptodonts but has a three-layered structure composed
of two layers of non-Haversian compact bone enclosing a central layer of
primary and secondary osteons. This internal organization is distinct from
other Astegotheriini of comparable age, but similar to Dasypus. The 3D
reconstruction of Riostegotherium yanei revealed two patterns of internal
organization. Pattern 1 of movable osteoderm is composed of large remodeled
areas at the base and a more compact bone at the tongue; in Pattern 2 (both
movable and buckler), the internal cavities are much smaller, more
numerous, and more interconnected to each other. In one buckler osteoderm,
the cavities are organized somewhat radially with a compact central region
(Pattern 1). Pattern 1 of both movable and buckler osteoderms resemble that
of Dasypus.


====

Espen M. Knutsen & Emma Oerlemans (2019)
The last dicynodont? Re-assessing the taxonomic and temporal relationships
of a contentious Australian fossil.
Gondwana Research (advance online publication)
doi: https://urldefense.proofpoint.com/v2/url?u=https-3A__doi.org_10.1016_j.gr.2019.07.011&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=owhrj2TMz-VYXRD2uuIQw9x4unEz1eiKjsriDIo8oDE&e= 
https://urldefense.proofpoint.com/v2/url?u=https-3A__www.sciencedirect.com_science_article_abs_pii_S1342937X19302254&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=NohkCg3H5E9wgwz5uphuJL31J8a5cL5eoeWWYAM4fKU&e= 

Highlights
Archives, geochemistry and anatomy reunite fossil specimens after 100 years.
Cretaceous dicynodont material now referred to Late Cenozoic mammalian
megafauna.
No evidence for a post-Triassic ghost lineage of Cretaceous dicynodonts

Abstract
Dicynodonts, a lineage of non-mammalian therapsids, who's derived taxa
evolved edentulous beaked jaws sporting a pair of caniniform tusks,
dominated the herbivorous terrestrial vertebrate fauna for much of the
Permian and Triassic periods. Long assumed to have met their demise during
the end-Triassic extinction event, the discovery of a fragmentary possible
dicynodont in Cretaceous rocks in Queensland Australia, potentially
extended the longevity of the lineage by nearly 100 million years. This
study reassesses the geological, anatomical and historical aspects of this
specimen through museum archival research, detrital zircon geochronology,
trace element analysis and x-ray synchrotron microtomography, and present
new knowledge regarding its temporal, geographical and biological origins,
supporting a late Cenozoic (Pliocene-Pleistocene) mammalian megafaunal
affinity for the specimen, resulting in a lack of evidence for
post-Triassic survival of dicynodonts.

=======

=======

Notice of New Applications to the Commission (Cases 3797–3812)
Bulletin of International Commission on Zoological Nomenclature 76: 98-99
https://urldefense.proofpoint.com/v2/url?u=https-3A__doi.org_10.21805_bzn.v76.a030&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=ldLDovMOwyTdBocNPEnt6NYBWo0Q5D0O1WmAhTXxQ4k&e= 
https://urldefense.proofpoint.com/v2/url?u=https-3A__bioone.org_journals_The-2DBulletin-2Dof-2DZoological-2DNomenclature_volume-2D76_issue-2D1_bzn.v76.a030_Notice-2Dof-2DNew-2DApplications-2Dto-2Dthe-2DCommission-2DCases-2D37973812_10.21805_bzn.v76.a030.short&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=_CA3w3ZaerUUSKfB8e5RfzHnXS7fjM5SR2ZAlyen0Wo&e= 

Free pdf:
https://urldefense.proofpoint.com/v2/url?u=https-3A__biotaxa.org_bzn_article_view_56367_50379&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=zK3oMEPjsNUllGDPkVYKuQ9CyHVucdczvSRSDtsDEZg&e= 

====

Only an abstract for now, full paper to come later:

Michelle C. Mekarski,  Stephanie E. Pierce and   Michael W. Caldwell (2019)
Spatiotemporal Distributions of Non-ophidian Ophidiomorphs, with
implications for their origin, radiation, and extinction.
Frontiers in Earth Science (abstract only)
doi: 10.3389/feart.2019.00245
https://urldefense.proofpoint.com/v2/url?u=https-3A__www.frontiersin.org_articles_10.3389_feart.2019.00245_abstract&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=MJ9LsbZmmRd0x_plGTOBbVA9RwLZ9ijI4HOu5ph2o9g&s=1dU-BXKRahcF7V5g0hx7cwM59dddPARqCkGJ1-dCBG0&e= 


Non-ophidian ophidiomorphs, colloquially referred to as 'dolichosaurs,' are
small-bodied aquatic lizards that lived in shallow seaways, rivers, and
reef environments during the Late Cretaceous. Preservational, geographic,
and taphonomic biases in this group make trends in biodiversity difficult
to assess. This is exemplified by the fact that the majority of the
described species are monotypic and known only from single specimens,
imparting very little information on morphological or spatial variation.
Here we present a revision of the spatial and temporal distributions of
non-ophidian ophidiomorph lizards ('dolichosaurs') from Cretaceous
sediments worldwide. The fossil record of dolichosaurs begins in the
Valanginian (Early Cretaceous). The late Early Cretaceous records are
sparse but suggest a wide geographic distribution spanning the Tethys and
Western Pacific. This is followed by a dense Cenomanian record from Tethyan
and British deposits, and rarer specimens from North America. Though there
is a substantial drop in the number of specimens recorded from the
Turonian-Maastrichtian, these rare occurrences represent the largest
geographical distribution of dolichosaurs: spanning Europe, North America,
and South America before going extinct during the end-Cretaceous mass
extinction. These occurrences indicate that ophidiomorphs most likely
originated in the Jurassic Tethys and continued to radiate spatially and
phylogenetically until the end of the Mesozoic, showing much more
temporally and environmentally diverse patterns than previously indicated.

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