[dinosaur] Hassiacoscutum, new chroniosuchian from Late Permian of Germany

Ben Creisler
[email protected]

A new paper:

Hassiacoscutum munki, gen. et sp. nov.


Florian Witzmann, Hans-Dieter Sues, Christian F. Kammerer & Jörg Fröbisch
(2019)
A new bystrowianid from the late Permian of Germany: First record of a
Permian chroniosuchian (Tetrapoda) outside Russia and China.
Journal of Vertebrate Paleontology Article: e1667366
doi: https://urldefense.proofpoint.com/v2/url?u=https-3A__doi.org_10.1080_02724634.2019.1667366&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=XQJNmJLZsakKYoHd6oZPZlQSklktQMOAxAnkYo8_ERY&s=OwcxluU57WjJDqTXLxUniZJzzDyAFWAlHK1zC7Iy5Dw&e= 
https://urldefense.proofpoint.com/v2/url?u=https-3A__www.tandfonline.com_doi_full_10.1080_02724634.2019.1667366&d=DwIFaQ&c=clK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=Ry_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=XQJNmJLZsakKYoHd6oZPZlQSklktQMOAxAnkYo8_ERY&s=6rrKTKPXZTU8P6drH9-iBbaTalAmKuAzO0xyTHco_10&e= 


An isolated dorsal osteoderm of a chroniosuchian from a late Permian
fissure filling in the lower Zechstein (Z1) of central Germany represents
the first Permian-age record of this enigmatic tetrapod clade outside
Russia and China. Based on a number of features, the specimen is designated
the holotype of a new taxon, Hassiacoscutum munki, and referred to the
chroniosuchian clade Bystrowianidae. Phylogenetic analysis based on a
recently published data set with addition of Hassiacoscutum and other
recently described bystrowianids (Yumenerpeton, Jiyuanitectum, and
Vyushkoviana) found good support for the clade Chroniosuchia and its
subclades Chroniosuchidae and Bystrowianidae, but with little internal
resolution for the latter. A second phylogenetic analysis of all known
bystrowianid genera based solely on osteoderm characters yielded a
well-supported and well-resolved clade Bystrowianidae with two subclades:
(Yumenerpeton + (Synesuchus + Bystrowiella)) and (Jiyuanitectum +
(Bystrowiana + (Hassiacoscutum + (Vyushkoviana + (Dromotectum +
Axitectum))))). Hassiacoscutum provides a paleobiogeographic link to other
middle and late Permian assemblages of Laurasian continental tetrapods. To
date, chroniosuchians are known only from Laurasia, and their apparent
absence in Gondwana hints at faunal provincialism across Pangaea.

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