Re: [dinosaur] Juvenile enantiornithines with immature feathers (free pdf)
Ben Creisler <[email protected]> Fri, 3 Jan 2020 13:22:11 -0800
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--00000000000009fa0f059b42e8ed Content-Type: text/plain; charset="UTF-8" Content-Transfer-Encoding: Quoted-printable The paper is now out in final form with new urls and a 2020 date:=0A=0AJing= mai K. O=E2=80=99CONNOR, Amanda FALK, WANG Min & ZHENG Xiao-Ting (2020)=0AF= irst report of immature feathers in juvenile enantiornithines from the=0AEa= rly Cretaceous Jehol avifauna.=0AVertebrate PalAsiatica 59(1): 24-44=0ADOI:= 10.19615/j.cnki.1000-3118.190823=0Ahttps://urldefense.proofpoint.com/v2/ur= l?u=3Dhttp-3A__www.ivpp.cas.cn_cbw_gjzdwxb_xbwzxz_201908_t20190823-5F536824= 5.html&d=3DDwIFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO= 4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8AhsC0Ekz5PRkA420zQ6Str8= Do1FL-cco-k&s=3DCjmHaBjWZoYb1cIC7k0SdheQ-cS_5LsYFNKaCacKkvY&e=3D=20=0A=0AFr= ee pdf:=0Ahttps://urldefense.proofpoint.com/v2/url?u=3Dhttp-3A__www.ivpp.ca= s.cn_cbw_gjzdwxb_xbwzxz_201908_P020200103352922223684.pdf&d=3DDwIFaQ&c=3Dcl= K7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn= 5g4z1CYJgFW9SI&m=3D_r1oI4X-8AhsC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&s=3De7pJJgCo= rLaBB3VIECPAlQ3N1Dk0ZbATafayRxV_jdU&e=3D=20=0A=0AOn Fri, Aug 23, 2019 at 9:= 47 AM Ben Creisler <[email protected]> wrote:=0A=0A>=0A> Ben Creisler=0A>= [email protected]=0A>=0A> A new paper with free pdf:=0A>=0A> Jingmai K. = O=E2=80=99CONNOR, Amanda FALK, WANG Min & ZHENG Xiao-Ting (2019)=0A> First = report of immature feathers in juvenile enantiornithines from the=0A> Early= Cretaceous Jehol avifauna.=0A> Vertebrata PalAsiatica (advance online publ= ication)=0A> DOI: 10.19615/j.cnki.1000-3118.190823=0A> https://urldefense.p= roofpoint.com/v2/url?u=3Dhttp-3A__www.ivpp.ac.cn_cbw_gjzdwxb_pressonline_20= 1908_t20190823-5F5368245.html&d=3DDwIFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN= 0H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8= AhsC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&s=3DjvEr5ASKkmS5Galmq44KWxSa0Q3qajkB8jfn= bxm_9FQ&e=3D=20=0A>=0A> Free pdf:=0A>=0A> https://urldefense.proofpoint.com= /v2/url?u=3Dhttp-3A__www.ivpp.ac.cn_cbw_gjzdwxb_pressonline_201908_P0201908= 23340535252530.pdf&d=3DDwIFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_= gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8AhsC0Ekz5PR= kA420zQ6Str8Do1FL-cco-k&s=3DF8YvC4tgKp1P2UzK-nig0CWmCNhrVESdVG8uW3oqkWU&e= =3D=20=0A>=0A> Molting--the process replacing one plumage with another--is = a critically=0A> important biological function in Aves. This process annual= ly replaces the=0A> feather coat, damaged by normal wear and tear, produces= ontogenetic changes=0A> in feathering, and produces alternate breeding plu= mages associated with=0A> reproductive activity in adults. Immature, growin= g feathers are encased in=0A> a keratinous sheath, giving them a narrow, tu= bular, and featureless=0A> appearance. The complete loss of the sheath indi= cates the feather is=0A> mature. Despite the wealth of integumentary data p= ublished from the Jehol=0A> Biota, immature feathers have never been defini= tively reported, although=0A> they may potentially be preserved in a juveni= le specimen of the non-avian=0A> oviraptorosaur theropod dinosaur Similicau= dipteryx from the 120 Ma=0A> Jiufotang Formation. A developing feather has = been reported in a 99 Ma=0A> enantiornithine neonate preserved in Burmese a= mber, in which=0A> three-dimensional preservation makes interpretations of = integumentary=0A> structures more straightforward. Here we report on probab= le immature=0A> feathers in four juvenile enantiornithines (Aves: Ornithoth= oraces) from the=0A> Jehol Group. As observed in developing feathers in ext= ant birds, the=0A> purported immature fossil feathers appear proximally nar= row and featureless=0A> with barbs protruding only distally. Based on our o= bservations, we suggest=0A> that similar-appearing feather structures prese= rved on the manus and=0A> tibiotarsus in the holotype of the enantiornithin= e Cruralispennia=0A> multidonta may alternatively be interpreted as immatur= e feathers. The=0A> presence of immature feathers in combination with sexua= lly dimorphic=0A> ornamental feathers in juvenile enantiornithines suggests= the complex=0A> molting patterns of Neornithes, in which such ornaments on= ly appear after=0A> several years (following several molts) when reproducti= ve activity is=0A> achieved, are limited to a subset of crownward avians.= =0A>=0A>=0A> <https://urldefense.proofpoint.com/v2/url?u=3Dhttp-3A__www.avg= .com_email-2Dsignature-3Futm-5Fmedium-3Demail-26utm-5Fsource-3Dlink-26utm-5= Fcampaign-3Dsig-2Demail-26utm-5Fcontent-3Dwebmail&d=3DDwIFaQ&c=3DclK7kQUTWt= AVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJ= gFW9SI&m=3D_r1oI4X-8AhsC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&s=3DqYdQ8Vr61nDQOpds= vwB8zgxgjdDy8UOzkmJOtJob7-o&e=3D > Virus-free.=0A> www.avg.com=0A> <https:/= /urldefense.proofpoint.com/v2/url?u=3Dhttp-3A__www.avg.com_email-2Dsignatur= e-3Futm-5Fmedium-3Demail-26utm-5Fsource-3Dlink-26utm-5Fcampaign-3Dsig-2Dema= il-26utm-5Fcontent-3Dwebmail&d=3DDwIFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0= H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8A= hsC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&s=3DqYdQ8Vr61nDQOpdsvwB8zgxgjdDy8UOzkmJOt= Job7-o&e=3D >=0A> <#m_1692498144654020603_DAB4FAD8-2DD7-40BB-A1B8-4E2AA1F9F= DF2>=0A>=0A= --00000000000009fa0f059b42e8ed Content-Type: text/html; charset="UTF-8" Content-Transfer-Encoding: Quoted-printable <div dir=3D"ltr">The paper is now out in final form with new urls and a 202= 0 date:<div><br></div><div>Jingmai K. O=E2=80=99CONNOR, Amanda FALK, WANG M= in & ZHENG Xiao-Ting (2020) <br>First report of immature feathers in ju= venile enantiornithines from the Early Cretaceous Jehol avifauna.<br>Verteb= rate PalAsiatica 59(1): 24-44 =C2=A0<br>DOI: 10.19615/j.cnki.1000-3118.1908= 23<br><a href=3D"https://urldefense.proofpoint.com/v2/url?u=3Dhttp-3A__www.= ivpp.cas.cn_cbw_gjzdwxb_xbwzxz_201908_t20190823-5F5368245.html&d=3DDwMF= aQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUmG= of_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8AhsC0Ekz5PRkA420zQ6Str8Do1= FL-cco-k&s=3DCjmHaBjWZoYb1cIC7k0SdheQ-cS_5LsYFNKaCacKkvY&e=3D">http= ://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/201908/t20190823_5368245.html</a><br>= <br>Free pdf:<br><a href=3D"https://urldefense.proofpoint.com/v2/url?u=3Dht= tp-3A__www.ivpp.cas.cn_cbw_gjzdwxb_xbwzxz_201908_P020200103352922223684.pdf= &d=3DDwMFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r= =3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8AhsC0Ekz5PR= kA420zQ6Str8Do1FL-cco-k&s=3De7pJJgCorLaBB3VIECPAlQ3N1Dk0ZbATafayRxV_jdU= &e=3D">http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/201908/P02020010335292= 2223684.pdf</a><br></div></div><br><div class=3D"gmail_quote"><div dir=3D"l= tr" class=3D"gmail_attr">On Fri, Aug 23, 2019 at 9:47 AM Ben Creisler <<= a href=3D"mailto:[email protected]">[email protected]</a>> wrote:<br= ></div><blockquote class=3D"gmail_quote" style=3D"margin:0px 0px 0px 0.8ex;= border-left:1px solid rgb(204,204,204);padding-left:1ex"><div dir=3D"ltr"><= br><div>Ben Creisler</div><div><a href=3D"mailto:[email protected]" targe= t=3D"_blank">[email protected]</a></div><div><br></div><div>A new paper w= ith free pdf:</div><div><br></div><div>Jingmai K. O=E2=80=99CONNOR, Amanda = FALK, WANG Min & ZHENG Xiao-Ting (2019)<br>First report of immature fea= thers in juvenile enantiornithines from the Early Cretaceous Jehol avifauna= .<br>Vertebrata PalAsiatica (advance online publication)<br>DOI: 10.19615/j= .cnki.1000-3118.190823<br><a href=3D"https://urldefense.proofpoint.com/v2/u= rl?u=3Dhttp-3A__www.ivpp.ac.cn_cbw_gjzdwxb_pressonline_201908_t20190823-5F5= 368245.html&d=3DDwMFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc= _gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8Ah= sC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&s=3DjvEr5ASKkmS5Galmq44KWxSa0Q3qajkB8j= fnbxm_9FQ&e=3D" target=3D"_blank">http://www.ivpp.ac.cn/cbw/gjzdwxb/pre= ssonline/201908/t20190823_5368245.html</a><br><br>Free pdf:<br><a href=3D"h= ttps://urldefense.proofpoint.com/v2/url?u=3Dhttp-3A__www.ivpp.ac.cn_cbw_gjz= dwxb_pressonline_201908_P020190823340535252530.pdf&d=3DDwMFaQ&c=3Dc= lK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgec= RCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X-8AhsC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&= ;s=3DF8YvC4tgKp1P2UzK-nig0CWmCNhrVESdVG8uW3oqkWU&e=3D" target=3D"_blank= ">http://www.ivpp.ac.cn/cbw/gjzdwxb/pressonline/201908/P0201908233405352525= 30.pdf</a><br><br>Molting--the process replacing one plumage with another--= is a critically important biological function in Aves. This process annuall= y replaces the feather coat, damaged by normal wear and tear, produces onto= genetic changes in feathering, and produces alternate breeding plumages ass= ociated with reproductive activity in adults. Immature, growing feathers ar= e encased in a keratinous sheath, giving them a narrow, tubular, and featur= eless appearance. The complete loss of the sheath indicates the feather is = mature. Despite the wealth of integumentary data published from the Jehol B= iota, immature feathers have never been definitively reported, although the= y may potentially be preserved in a juvenile specimen of the non-avian ovir= aptorosaur theropod dinosaur Similicaudipteryx from the 120 Ma Jiufotang Fo= rmation. A developing feather has been reported in a 99 Ma enantiornithine = neonate preserved in Burmese amber, in which three-dimensional preservation= makes interpretations of integumentary structures more straightforward. He= re we report on probable immature feathers in four juvenile enantiornithine= s (Aves: Ornithothoraces) from the Jehol Group. As observed in developing f= eathers in extant birds, the purported immature fossil feathers appear prox= imally narrow and featureless with barbs protruding only distally. Based on= our observations, we suggest that similar-appearing feather structures pre= served on the manus and tibiotarsus in the holotype of the enantiornithine = Cruralispennia multidonta may alternatively be interpreted as immature feat= hers. The presence of immature feathers in combination with sexually dimorp= hic ornamental feathers in juvenile enantiornithines suggests the complex m= olting patterns of Neornithes, in which such ornaments only appear after se= veral years (following several molts) when reproductive activity is achieve= d, are limited to a subset of crownward avians.<br></div></div><div id=3D"g= mail-m_1692498144654020603DAB4FAD8-2DD7-40BB-A1B8-4E2AA1F9FDF2"><br>=0A<tab= le style=3D"border-top:1px solid rgb(211,212,222)">=0A <tbody><tr>=0A = <td style=3D"width:55px;padding-top:13px"><a href=3D"https://urldefense.p= roofpoint.com/v2/url?u=3Dhttp-3A__www.avg.com_email-2Dsignature-3Futm-5Fmed= ium-3Demail-26utm-5Fsource-3Dlink-26utm-5Fcampaign-3Dsig-2Demail-26utm-5Fco= ntent-3Dwebmail&d=3DDwMFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7C= Sfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3D_r1oI4X= -8AhsC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&s=3DqYdQ8Vr61nDQOpdsvwB8zgxgjdDy8U= OzkmJOtJob7-o&e=3D" target=3D"_blank"><img src=3D"https://ipmcdn.avast.= com/images/icons/icon-envelope-tick-green-avg-v1.png" alt=3D"" width=3D"46"= height=3D"29" style=3D"width: 46px; height: 29px;"></a></td>=0A <td style= =3D"width:470px;padding-top:12px;color:rgb(65,66,78);font-size:13px;font-fa= mily:Arial,Helvetica,sans-serif;line-height:18px">Virus-free. <a href=3D"ht= tps://urldefense.proofpoint.com/v2/url?u=3Dhttp-3A__www.avg.com_email-2Dsig= nature-3Futm-5Fmedium-3Demail-26utm-5Fsource-3Dlink-26utm-5Fcampaign-3Dsig-= 2Demail-26utm-5Fcontent-3Dwebmail&d=3DDwMFaQ&c=3DclK7kQUTWtAVEOVIgv= i0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9= SI&m=3D_r1oI4X-8AhsC0Ekz5PRkA420zQ6Str8Do1FL-cco-k&s=3DqYdQ8Vr61nDQ= OpdsvwB8zgxgjdDy8UOzkmJOtJob7-o&e=3D" style=3D"color:rgb(68,83,234)" ta= rget=3D"_blank">www.avg.com</a>=0A </td>=0A </tr>=0A</tbody></table><a hre= f=3D"#m_1692498144654020603_DAB4FAD8-2DD7-40BB-A1B8-4E2AA1F9FDF2" width=3D"= 1" height=3D"1"></a></div>=0A</blockquote></div>=0A= --00000000000009fa0f059b42e8ed--