[dinosaur] Yunyangosaurus, new megalosauroid from Middle Jurassic of China (free pdf)

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Ben [email protected]=0A=0A=0AA new paper with free pdf:=0A=0A=
=0AYunyangosaurus puanensis gen. et sp. nov.=0A=0AHui Dai, Roger Benson, Xu=
feng Hu, Qingyu Ma, Chao Tan, Ning Li, Ming Xiao,=0AHaiqian Hu, Yuxuan Zhou=
, Zhaoying Wei, Feng Zhang, Shan Jiang, Deliang Li,=0AGuangzhao Peng, Yilun=
 Yu & Xing Xu (2020)=0AA new possible megalosauroid theropod from the Middl=
e Jurassic Xintiangou=0AFormation of Chongqing, People=E2=80=99s Republic o=
f China and its implication for=0Aearly tetanuran evolution.=0AScientific R=
eports 10, Article number: 139=0Adoi: https://urldefense.proofpoint.com/v2/=
url?u=3Dhttps-3A__doi.org_10.1038_s41598-2D019-2D56959-2Dx&d=3DDwIFaQ&c=3Dc=
lK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKH=
n5g4z1CYJgFW9SI&m=3DATPVWMsBcgPIIx4IiYbFM6qvHaXXlfqWkSgRYZMCDfs&s=3DsmlqSRL=
t4hbXhk5AYy5U7nwIp37NXGmbHsAbYvwc1h0&e=3D=20=0Ahttps://urldefense.proofpoin=
t.com/v2/url?u=3Dhttps-3A__www.nature.com_articles_s41598-2D019-2D56959-2Dx=
&d=3DDwIFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUm=
Gof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3DATPVWMsBcgPIIx4IiYbFM6qvHaXXlfqWkSgRYZ=
MCDfs&s=3DQYGOjsBBUYacrlJ-g0L8Vt94_xS0McKccEMzvwnQd6E&e=3D=20=0A=0AFree pdf=
:=0Ahttps://urldefense.proofpoint.com/v2/url?u=3Dhttps-3A__www.nature.com_a=
rticles_s41598-2D019-2D56959-2Dx.pdf&d=3DDwIFaQ&c=3DclK7kQUTWtAVEOVIgvi0NU5=
BOUHhpN0H8p7CSfnc_gI&r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&m=3DAT=
PVWMsBcgPIIx4IiYbFM6qvHaXXlfqWkSgRYZMCDfs&s=3DyfThNA6sDHBOSKTGh_YEamQK3eTtR=
zIWaN0FLzkr7F0&e=3D=20=0A=0A=0ATetanurae is a special group of theropod din=
osaurs that originated by the=0Alate Early Jurassic. It includes several ea=
rly-diverging groups of=0Agenerally large-bodied predators (megalosauroids,=
 allosauroids,=0Atyrannosauroid coelurosaurs) as well as morphologically di=
sparate=0Asmall-bodied coelurosaurs, including birds. Aspects of the evolut=
ionary=0Ahistory of tetanurans remain contested, including the topology of =
their=0Adeep phylogenetic divergences (among Megalosauroidea, Allosauroidea=
 and=0ACoelurosauria). We report a new theropod, Yunyangosaurus puanensis g=
en. et=0Asp. nov., based on a fragmentary specimen recovered from the Middl=
e=0AJurassic Xintiangou Formation of Chongqing, southwestern China. It shar=
es=0Aseveral features uniquely with some megalosauroids (the clade of=0Ameg=
alosaurids + spinosaurids + piatnitzkysaurids), such as prominent rims=0Aar=
ound the anterior articular surfaces of cervical centra and bifurcated=0Aan=
terior dorsal neural spines (present in piatnitzkysaurids). Nevertheless,=
=0Ait also shows several features that are rare or absent among megalosauro=
ids=0Aand more crownward tetanurans, including prominent spinopostyzgopophy=
seal=0Alaminae (also present in non-tetanurans and metriacanthosaurid=0Aall=
osauroids), flat anterior articular surfaces of the cervical centra=0A(also=
 present in piatnitzkysaurids and some earlier-diverging tetanurans),=0Aand=
 the presence of a posterior pneumatic foramen or fossa (absent in most=0At=
etanurans, but sporadically present in some cervical vertebrae of=0Apiatnit=
zkysaurids). Yunyangosaurus therefore presents a combination of=0Aderived a=
nd apparently primitive character states that are not seen in=0Aother thero=
pods. This suggests that patterns of morphological evolution=0Aassociated w=
ith deep tetanuran divergences were more complex than currently=0Arecognize=
d, with implications for understanding the character evolution in=0Atheropo=
ds.=0A=

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<div dir=3D"ltr"><div><br></div>Ben Creisler<div><a href=3D"mailto:bcreisle=
[email protected]">[email protected]</a></div><div><br></div><div><br></div><di=
v>A new paper with free pdf:<br><div><br></div><div><br></div><div>Yunyango=
saurus puanensis gen. et sp. nov.=C2=A0</div><div><br>Hui Dai, Roger Benson=
, Xufeng Hu, Qingyu Ma, Chao Tan, Ning Li, Ming Xiao, Haiqian Hu, Yuxuan Zh=
ou, Zhaoying Wei, Feng Zhang, Shan Jiang, Deliang Li, Guangzhao Peng, Yilun=
 Yu &amp; Xing Xu (2020)<br>A new possible megalosauroid theropod from the =
Middle Jurassic Xintiangou Formation of Chongqing, People=E2=80=99s Republi=
c of China and its implication for early tetanuran evolution.<br>Scientific=
 Reports 10, Article number: 139<br>doi: <a href=3D"https://urldefense.proo=
fpoint.com/v2/url?u=3Dhttps-3A__doi.org_10.1038_s41598-2D019-2D56959-2Dx&am=
p;d=3DDwMFaQ&amp;c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&amp;r=3DRy=
_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&amp;m=3DATPVWMsBcgPIIx4IiYbFM6qvH=
aXXlfqWkSgRYZMCDfs&amp;s=3DsmlqSRLt4hbXhk5AYy5U7nwIp37NXGmbHsAbYvwc1h0&amp;=
e=3D">https://doi.org/10.1038/s41598-019-56959-x</a><br><a href=3D"https://=
urldefense.proofpoint.com/v2/url?u=3Dhttps-3A__www.nature.com_articles_s415=
98-2D019-2D56959-2Dx&amp;d=3DDwMFaQ&amp;c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0=
H8p7CSfnc_gI&amp;r=3DRy_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&amp;m=3DAT=
PVWMsBcgPIIx4IiYbFM6qvHaXXlfqWkSgRYZMCDfs&amp;s=3DQYGOjsBBUYacrlJ-g0L8Vt94_=
xS0McKccEMzvwnQd6E&amp;e=3D">https://www.nature.com/articles/s41598-019-569=
59-x</a><br><br>Free pdf:<br><a href=3D"https://urldefense.proofpoint.com/v=
2/url?u=3Dhttps-3A__www.nature.com_articles_s41598-2D019-2D56959-2Dx.pdf&am=
p;d=3DDwMFaQ&amp;c=3DclK7kQUTWtAVEOVIgvi0NU5BOUHhpN0H8p7CSfnc_gI&amp;r=3DRy=
_mO4IFaUmGof_Yl9MyZgecRCKHn5g4z1CYJgFW9SI&amp;m=3DATPVWMsBcgPIIx4IiYbFM6qvH=
aXXlfqWkSgRYZMCDfs&amp;s=3DyfThNA6sDHBOSKTGh_YEamQK3eTtRzIWaN0FLzkr7F0&amp;=
e=3D">https://www.nature.com/articles/s41598-019-56959-x.pdf</a><br><br><br=
>Tetanurae is a special group of theropod dinosaurs that originated by the =
late Early Jurassic. It includes several early-diverging groups of generall=
y large-bodied predators (megalosauroids, allosauroids, tyrannosauroid coel=
urosaurs) as well as morphologically disparate small-bodied coelurosaurs, i=
ncluding birds. Aspects of the evolutionary history of tetanurans remain co=
ntested, including the topology of their deep phylogenetic divergences (amo=
ng Megalosauroidea, Allosauroidea and Coelurosauria). We report a new thero=
pod, Yunyangosaurus puanensis gen. et sp. nov., based on a fragmentary spec=
imen recovered from the Middle Jurassic Xintiangou Formation of Chongqing, =
southwestern China. It shares several features uniquely with some megalosau=
roids (the clade of megalosaurids=E2=80=89+=E2=80=89spinosaurids=E2=80=89+=
=E2=80=89piatnitzkysaurids), such as prominent rims around the anterior art=
icular surfaces of cervical centra and bifurcated anterior dorsal neural sp=
ines (present in piatnitzkysaurids). Nevertheless, it also shows several fe=
atures that are rare or absent among megalosauroids and more crownward teta=
nurans, including prominent spinopostyzgopophyseal laminae (also present in=
 non-tetanurans and metriacanthosaurid allosauroids), flat anterior articul=
ar surfaces of the cervical centra (also present in piatnitzkysaurids and s=
ome earlier-diverging tetanurans), and the presence of a posterior pneumati=
c foramen or fossa (absent in most tetanurans, but sporadically present in =
some cervical vertebrae of piatnitzkysaurids). Yunyangosaurus therefore pre=
sents a combination of derived and apparently primitive character states th=
at are not seen in other theropods. This suggests that patterns of morpholo=
gical evolution associated with deep tetanuran divergences were more comple=
x than currently recognized, with implications for understanding the charac=
ter evolution in theropods.<br><br></div></div></div>=0A=

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